1. School of Geography and the Environment, Univ. of Oxford, South Parks Rd., Oxford OX1 3QY, U.K. (corresponding author).
2. School of Geography and the Environment, Univ. of Oxford, South Parks Rd., Oxford OX1 3QY, U.K.
3. Centre for Ecology and Hydrology (CEH), Benson Ln., Crowmarsh Gifford, Wallingford OX10 8BB, U.K.
4. Professor, School of Geography and the Environment, Univ. of Oxford, South Parks Rd., Oxford OX1 3QY, U.K.