1. Center for Computational Biology, Flatiron Institute, New York, NY 10010, U.S.A.
2. Center for Computational Biology, Flatiron Institute, New York, NY 10010, U.S.A., and Physics and Astronomy Department, Rutgers University, New Brunswick, NJ 08901, U.S.A.
3. Center for Computational Biology, Flatiron Institute, New York, NY 10010, U.S.A., and NYU Langone Medical Center, New York 10016, U.S.A.