1. Evolution and Ecology Research Centre, University of New South Wales, Sydney, NSW 2052, Australia, Sydney Institute of Marine Science, Chowder Bay Road, Mosman, NSW 2088, Australia;, Email: james.smith@unsw.edu.au
2. Centre for Coastal Biogeochemistry, Southern Cross University, East Lismore, NSW 2480, Australia
3. Evolution and Ecology Research Centre, University of New South Wales, Sydney, NSW 2052, Australia, Sydney Institute of Marine Science, Chowder Bay Road, Mosman, NSW 2088, Australia, Port Stephens Fisheries Institute, New South
Wales Department of Primary Industries, Taylors Beach Rd, Taylors Beach, New South Wales 2316, Australia