1. Evolution and Ecology Research Centre and School of Biological, Earth, and Environmental Sciences, University of New South Wales, Sydney, NSW, 2052, Australia
2. Northwest Atlantic Fisheries Centre, Fisheries and Oceans Canada, 80 East White Hills, P.O. Box 5667, St. John’s, NL A1C 5X1, Canada
3. Sydney Institute of Marine Science, Chowder Bay Road, Mosman, NSW 2088, Australia