1. School of Environmental Sciences University of Liverpool Liverpool UK
2. Centre d'Etudes Biologiques de Chizé Centre National de la Recherche Scientifique Villiers en Bois France
3. Institute of Infection, Veterinary and Ecological Sciences University of Liverpool Liverpool UK
4. School of Biosciences University of Sheffield Sheffield UK
5. Centre d'Ecologie et des Sciences de la Conservation (UMR 7204) Muséum National d'Histoire Naturelle Paris France