1. Department of Horticultural Science University of Minnesota St. Paul MN 55108 USA
2. Current affiliation: Department of Agronomy and Plant Genetics University of Minnesota St. Paul MN 55108 USA
3. Department of Plant and Microbial Biology University of Minnesota St. Paul MN 55108 USA
4. Department of Plant Pathology University of Minnesota St. Paul MN 55108 USA