1. CAS Key Laboratory of Tropical Forest Ecology Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Mengla Yunnan666303China
2. Department of Crop and Environment Sciences Harper Adams University NewportTF10 8NBUnited Kingdom
3. Centre d'Ecologie Fonctionnelle et Evolutive UMR 5175 CNRS–University of Montpellier–University Paul–Valéry–EPHE–SupAgro Montpellier–INRA–IRD Montpellier34293France