1. Jiangxi Province Key Laboratory of Watershed Ecosystem Change and Biodiversity; Center for Watershed Ecology; Institute of Life Science and School of Life Sciences; Nanchang University; Nanchang 330031 China
2. Key Laboratory of Poyang Lake Environment and Resource Utilization; Ministry of Education; Nanchang University; Nanchang 330031 China
3. National Ecosystem Research Station of Jiangxi Poyang Lake Wetland; Nanchang 330038 China
4. Department of Ecology and Evolutionary Biology; University of Toronto; 25 Willcocks St. Toronto Ontario Canada M5S 3B2
5. The Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering; Institute of Biodiversity Science; Fudan University; Shanghai 200438 China