1. Peony International Research Institute, National Flower Engineering Research Centre, Key Laboratory for the Genetics and Breeding of Forest Trees and Ornamental Plants, College of Landscape Architecture Beijing Forestry University Beijing China
2. Beijing Advanced Innovation Center for Tree Breeding by Molecular Design, National Engineering Laboratory for Tree Breeding, Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants, Ministry of Education, College of Biological Sciences and Technology Beijing Forestry University Beijing China
3. Forestry Department of Shaanxi Province Xi'an Shaanxi China
4. School of Biology, Mitchell Building University of St Andrews St Andrews Fife UK