1. Dep. of Plant Sciences and Plant Pathology Montana State Univ. 119 Plant Biosciences Bozeman MT 59717‐3150 USA
2. Current address: Dep. of Research Centers Montana State Univ. Conrad MT 59425 USA
3. USDA–ARS, Washington State Univ. Pullman USA
4. Crop and Soil Sciences Washington State Univ. College of Agricultural Human and Natural Resource Sciences 291B, Johnson Hall Pullman WA 99164 USA
5. USDA–ARS Wheat Health, Genetics and Quality Research Unit Pullman WA 99164 USA