1. College of Resources and Environment Huazhong Agricultural University Wuhan China
2. Wuhan Botanical Garden Chinese Academy of Sciences Wuhan China
3. Department of Biology University of Fribourg Fribourg Switzerland
4. Division of Biological Sciences and Wildlife Biology University of Montana Missoula Montana USA
5. Department of Biology University of Konstanz Konstanz Germany
6. Zhejiang Provincial Key Laboratory of Plant Evolutionary Ecology and Conservation Taizhou University Taizhou China