1. Kunming Institute of Botany; Key Laboratory of Biodiversity and Biogeography, Chinese Academy of Sciences; Kunming, Yunnan, P.R. 650201 China
2. University of Chinese Academy of Sciences; Beijing 100039 P.R. China
3. Jishou University; Key Laboratory of Plant Resources Conservation and Utilization, College of Hunan Province; Jishou, Hunan 416000 P.R. China
4. Yunnan Normal University; Life Science School; Kunming, Yunnan 650031 P.R. China