1. School of Marine Sciences, Sun Yat-Sen University; Guangzhou China
2. Key Laboratory of Marine Resources and Coastal Engineering in Guangdong Province; Guangzhou China
3. School for Marine Science and Technology, University of Massachusetts-Dartmouth; New Bedford Massachusetts USA
4. International Center for Marine Studies, Shanghai Ocean University; Shanghai China
5. Department of Physical Oceanography; Woods Hole Oceanographic Institution; Woods Hole Massachusetts USA