1. Pearl River Fisheries Research Institute Chinese Academy of Fishery Science Guangzhou China
2. Key Laboratory of Aquatic Animal Immune Technology of Guangdong Province Guangzhou China
3. Guangzhou Scientific Observing and Experimental Station of National Fisheries Resources and Environment Guangzhou China
4. Experimental Station for Scientific Observation on Fishery Resources and Environment in the Middle and Lower Reaches of Pearl River Zhaoqing China