1. College of Fisheries, Huazhong Agricultural University; Freshwater Aquaculture Collaborative Innovation Center of Hubei Province; Wuhan 430070 China
2. Collaborative Innovation Center for Efficient and Health Production of Fisheries in Hunan Province, Key Laboratory of Health Aquaculture and Product Processing in Dongting Lake Area of Hunan Province; Hunan University of Arts and Science; Hunan Changde 415000 China