1. CAS Key Laboratory of Mountain Ecological Restoration and Bioresource UtilizationChengdu Institute of Biology, Chinese Academy of Sciences Chengdu 610041 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. Department of Plant sciencesQuaid‐i‐Azam University Islamabad Pakistan
4. State Key Laboratory of Systematic and Evolutionary BotanyInstitute of Botany, Chinese Academy of Sciences Beijing 100093 China
5. Department of BiologyUniversity of Florence Italy
6. Depratment of BotanyUniversity of Peshawar Peshawar Pakistan