1. Department of Anatomy, Faculty of Medicine; Toho University; Tokyo 143-8540 Japan
2. Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Agency; Saitama 332-0012 Japan
3. Department of Physiology, Faculty of Medicine; Toho University; Tokyo 143-8540 Japan
4. International institute for integrative sleep medicine (IIIS), Tsukuba University; Ibaraki 305-8575 Japan