1. Department of Neurobiology , Graduate School of Biomedical and Health Sciences, , 1-2-3 Kasumi, Minami-ku, Hiroshima City, Hiroshima 734-8553 , Japan
2. Hiroshima University , Graduate School of Biomedical and Health Sciences, , 1-2-3 Kasumi, Minami-ku, Hiroshima City, Hiroshima 734-8553 , Japan