1. Key Laboratory of Vertebrate Evolution and Human Origins of Chinese Academy of Sciences; Institute of Vertebrate Paleontology and Paleoanthropology Chinese Academy of Sciences; Beijing 100044 China
2. Department of Anthropology; University College London (UCL), 14 Taviton Street; London WC1H 0BW
3. Laboratorio de Evolución Humana, Departamento de Historia, Geografía y Comunicación, University of Burgos; Burgos 09001 Spain
4. Department of Anthropology; National Museum of Nature and Science, 4-1-1 Amakubo, Tsukuba-shi; Ibaraki 305-0005 Japan
5. Department of Geology; National Museum of Natural Science; Taichung 40453 Taiwan
6. State Key Laboratory of Genetic Engineering and Ministry of Education Key Laboratory of Contemporary Anthropology, School of Life Sciences, Fudan University; Shanghai 200433 China
7. Hominid Paleobiology Program; National Research Center on Human Evolution (CENIEH), Paseo Sierra de Atapuerca s/n. Burgos; 09002 Spain