1. Key Laboratory of Industrial Fermentation Microbiology of the Ministry of Education Tianjin Key Laboratory of Industrial Microbiology College of Biotechnology Tianjin University of Science and Technology National Engineering Laboratory for Industrial Enzymes Tianjin 300457 China
2. Luoyang BIO-Industry Technology Innovation Center Luoyang 471000 Henan China
3. Department of Applied Biological Chemistry Graduate School of Agricultural and Life Sciences The University of Tokyo Bunkyo Tokyo 113-8657 Japan