1. Key Laboratory of Karst Georesources and Environment Ministry of Education, Guizhou University Guiyang PR China
2. Institute for Forest Resources and Environment of Guizhou Guizhou University Guiyang PR China
3. College of Forestry Guizhou University Guiyang PR China
4. Forestry Bureau Tongren PR China
5. School of Geography & Environmental Science Guizhou Normal University Guiyang PR China