1. Idaho Department of Fish and Game; 600 South Walnut Street; Boise ID 83712 USA
2. Wildlife Biology Program; Department of Ecosystem and Conservation Sciences, W.A. Franke College of Forestry and Conservation, University of Montana; Missoula MT 59812 USA
3. Wildlife Biology Program; W.A. Franke College of Forestry and Conservation; University of Montana; Missoula MT 59812 USA
4. Laboratoire Biométrie et Biologie Évolutive; UMR-CNRS 5558, University Claude Bernard − Lyon I; 69622 Villeurbanne Cedex France