1. Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering, National Observations and Research Station for Wetland Ecosystems of the Yangtze Estuary, Institute of Biodiversity Science and Institute of Eco‐Chongming, School of Life Sciences Fudan University Shanghai China
2. Key Laboratory of the Ministry of Education for Coastal and Wetland Ecosystems, College of the Environment and Ecology Xiamen University Fujian China
3. Department of Biology and Biochemistry University of Houston Houston Texas USA
4. Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology and Centre for Invasion Biology, Institute of Biodiversity, School of Ecology and Environmental Science Yunnan University Kunming Yunnan China