1. Department of Ecology and Evolutionary Biology University of Michigan Ann Arbor Michigan48019USA
2. Department of Biological Sciences University of Notre Dame South Bend Indiana46556USA
3. Institut de recherche sur les forêts Université du Québec en Abitibi‐Témiscamingue Rouyn‐Noranda QuebecJ9X 5E4Canada
4. CAS Key Laboratory of Tropical Forest Ecology Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Kunming Yunnan China
5. Institute of Eastern‐Himalaya Biodiversity Research Dali University Dali Yunnan China