1. School of Biological Sciences, University of Portsmouth, Portsmouth PO12DY, UK
2. Institute of Arctic Biology, University of Alaska, Fairbanks, AK 99775, USA
3. Department of Biology, Norwegian University of Science and Technology, 7491 Trondheim, Norway
4. Center for Biodiversity Dynamics, Department of Biology, Norwegian University of Science and Technology, 7491 Trondheim, Norway
5. Centre for Ecological and Evolutionary Synthesis, Department of Biology, University of Oslo, PO Box 1066, 0316 Oslo, Norway