1. College of Biology and Agriculture Zunyi Normal College Zunyi 563002Guizhou China
2. Chinese Academy of Sciences South China Sea Institute of Oceanology Guangdong Provincial Key Laboratory of Applied Marine Biology, CAS Key Laboratory of Tropical Marine Bio‐resources and Ecology Guangzhou 510301 China
3. University of Chinese Academy of Sciences Beijing 100049 China
4. Engineering Research Center of Tropical and Subtropical Aquatic Ecological Engineering Ministry of Education Key Laboratory of Aquatic Eutrophication and Red Tide Prevention of Guangdong Higher Education Institutes Institute of Hydrobiology Jinan University West 601 Huangpu Avenue Tianhe District Guangzhou 510632 China
5. Special Fish Protection and Development and Utilization Engineering Research Center of North Guizhou Province Zunyi 563002Guizhou China