1. Key Laboratory of Vertebrate Evolution and Human Origins; Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences; Beijing China
2. CAS Center for Excellence in Life and Paleoenvironment; Beijing China
3. University of Chinese Academy of Sciences; Beijing China
4. Histoire Naturelle de l'Homme Préhistorique (HNHP, UMR 7194 CNRS); Tautavel France