1. Tunku Abdul Rahman University College Johor Branch Campus Segamat Malaysia
2. CNRS ‐ University of Corsica UMR 6134 ‐ SPE Natural Resources Project Vignola Route des Sanguinaires Ajaccio F‐20000 France
3. Institute of Biodiversity and Environmental Conservation Universiti Malaysia Sarawak Samarahan Sarawak 94300 Malaysia
4. Research Associate Harvard University Herbaria 22 Divinity Avenue Cambridge MA 02138 USA