1. Graduate School of Horticulture Chiba University 648 Matsudo, Matsudo Chiba 271‐8510 Japan
2. Gifu Academy of Forest Science and Culture 88 Sodai, Mino Gifu Japan
3. Department of Biology Daejeon University 62 Daehak‐ro, Dong‐gu Daejeon 34520 South Korea
4. Faculty of Agriculture Ryukoku University 1‐5 Yokotani, Seta Oe‐cho, Otsu Shiga 520‐2194 Japan
5. Institute for Advanced Biosciences Keio University 403‐1 Nipponkoku, Daihouji, Tsuruoka Yamagata 997‐0017 Japan
6. College of Liberal Arts and Sciences Chiba University 1‐33 Yayoi‐cho, Inage‐ku Chiba 263‐8522 Japan
7. Graduate School of Bioagricultural Sciences Nagoya University Furo‐cho, Chikusa Nagoya Japan
8. Sado Island Center for Ecological Sustainability, Niigata University 94‐2 Koda, Sado Niigata 952‐2206 Japan