1. Laboratory of Evolution and Biologic Diversity; UMR 5174 CNRS/UPS/ENFA; Narbonne Road PB 22687 31326 Castanet Tolosan Cedex France
2. Department of Biology; Faculty of Sciences of Bizerte; University of Carthage; Jarzouna Bizerte 7021 Carthage Tunisia
3. Laboratory of Materials, Molecules and Applications (LMMA); University of Carthage, IPEST; PB 51 La Marsa 2070 Tunisia
4. Laboratory of Bioactive Substances; Biotechnology Center in Borj-Cedria Technopol; PB 901 2050 Hammam-Lif Tunisia
5. Pharmacy Department; Faculty of Medicine; University of Antananarivo; PB 375 Antananarivo 101 Madagascar
6. Laboratory of Molecular Interactions and Chemical and Photochemical Reactivity; UMR CNRS 5623; Paul-Sabatier University; 118 Narbonne Road F-31062 Toulouse France