1. Graduate School of Life and Environmental Sciences; Osaka Prefecture University; 1-1 Gakuen-cho Naka-ku Sakai Osaka 599-8531 Japan
2. Laboratory of Plant Pathology and Biotechnology; Kochi University; 200 Otsu Monobe Nanko-ku Kochi 783-8502 Japan
3. Research Institute of Molecular Genetics; Kochi University; 200 Otsu Monobe Nanko-ku Kochi 783-8502 Japan
4. Faculty of Agriculture; Kagawa University; 2393 Ikenobe Miki-cho Kagawa 761-0795 Japan