1. Department of Biology Faculty of Science Toho University Chiba Japan
2. Agro‐biological Resource Sciences University of Tsukuba Tsukuba Japan
3. Department of Forest Molecular Genetics and Biotechnology Forestry and Forest Products Research InstituteForest Research and Management Organization Tsukuba Japan
4. Institute of Plant Science and Resources Okayama University Kurashiki Japan
5. Tohoku Agricultural Research CenterNARO Arai Fukushima Japan