1. Department of Biology, Biodiversity Unit Lund University Lund Sweden
2. Research Centre for Ecological changes, Organismal and Evolutionary Biology Research Program Faculty of Environmental and Biological Sciences University of Helsinki Helsinki Finland
3. Laboratory of Microbial Ecology UMR CNRS 5557 UMR INRA 1418 University Claude Bernard Lyon 1 Villeurbanne France
4. Helsinki Institute of Life Science University of Helsinki Helsinki Finland