1. School of Life Sciences; University of Warwick; Gibbet Hill, Coventry CV4 7AL U.K
2. School of Biosciences; University of Nottingham; Sutton Boningon Campus Sutton Bonington, Leicestershire LE12 5RD U.K
3. INRA Laboratoire d'Ecophysiologie des Plantes sous Stress Environnementaux (LEPSE); Montpellier France
4. Department of Biology; University of York; Heslington, York YO10 5DD U.K