1. Division of Biological Anthropology, Department of Archaeology & Anthropology; University of Cambridge; Cambridge CB2 3QG UK
2. Infection, Inflammation and Rheumatology Section; UCL Institute of Child Health; London WC1N 1EH UK
3. Human Origins and Palaeoenvironmental Research Group (HOPE), Department of Anthropology & Geography; Oxford Brookes University; Oxford OX3 0BP UK