1. Office of the Chief Director: Biosystematics Research & Biodiversity Collections; South African National Biodiversity Institute; Private Bag X101 Pretoria 0001 South Africa
2. H.G.W.J. Schweickerdt Herbarium; Department of Plant Science; University of Pretoria; Pretoria 0002 South Africa
3. Centre for Functional Ecology; Departamento de Ciências da Vida; Universidade de Coimbra; 3001-455 Coimbra Portugal
4. Department of Botany; Nelson Mandela Metropolitan University; P.O. Box 77000 Port Elizabeth 6031 South Africa