1. Brooklyn Botanic Garden,; 1000 Washington Avenue, Brooklyn, New York 11225, U.S.A.;,
2. Department of Plant Science,; Office of the Chief Director: Biosystematics Research and Biodiversity Collections, South African National Biodiversity Institute,; Private Bag X101, / Acocks Chair, H.G.W.J. Schweickerdt Herbarium, University of Pretoria, Pretoria,; Pretoria, South Africa 0001 0002 South Africa
3. Department of Plant Science,; H.G.W.J. Schweickerdt Herbarium, University of Pretoria, ; Pretoria, 0002 South Africa
4. The National Herbarium, Addis Ababa University,; P.O. Box Addis Ababa, 3434, Ethiopia
5. Royal Botanic Gardens, Kew, Herbarium, Library, Art and Archives,; Richmond, Surrey, TW9 3AB, U.K.