1. CERMAV; Univ. Grenoble Alpes; Grenoble F-38000 France
2. CERMAV; CNRS; Grenoble F-38000 France
3. Univ. Grenoble Alpes, LGP2; Grenoble F-38000 France
4. CNRS, LGP2; Grenoble F-38000 France
5. UMR IATE, UM2-CIRAD-INRA-SupAgro; 2 Pl Pierre Viala Montpellier 34070 France