1. Centre for Ecology and Conservation; College of Life and Environmental Sciences; University of Exeter; Tremough Campus Penryn TR10 9EZ UK
2. Department of Entomology; North Carolina State University; Gardner Hall Raleigh North Carolina 27695-7613
3. MaxNetAging School; Max Planck Institute for Demographic Research; Konrad-Zuse-Straße 1 18057 Rostock Germany