1. Department of Fish and Wildlife ConservationVirginia Polytechnic Institute and State University Blacksburg VA 24061 USA
2. Department of Natural SciencesMars Hill University Mars Hill NC 28754 USA
3. Wildlife Management DivisionNorth Carolina Wildlife Resources Commission Asheville NC 28803 USA
4. U.S. Geological SurveyVirginia Cooperative Fish and Wildlife Research Unit Blacksburg VA 24061 USA