1. Wildlife Conservation Research Unit Department of Zoology Recanati‐Kaplan Centre University of Oxford Tubney House Abingdon Road Tubney Abingdon Oxfordshire OX13 5QL U.K
2. NERC Biomolecular Analysis Facility Department of Animal and Plant Sciences University of Sheffield Sheffield S10 2TN U.K
3. Faculty of Science Department of Biology University of Putra Malaysia UPM 43400 Serdang Selangor Malaysia
4. Theoretical Biology Centre for Ecological and Evolutionary Studies University of Groningen PO Box 11103 9700 CC Groningen The Netherlands
5. Behavioural Ecology and Self‐Organization Centre for Ecological and Evolutionary Studies University of Groningen PO Box 11103 9700 CC Groningen The Netherlands