1. Stellenbosch Institute for Advanced Study, Stellenbosch 7600, South Africa
2. Department of Molecular Cell Physiology, BioCentrum Amsterdam, Faculty of biology, Vrije Universiteit, De Boelelaan 1087, Amsterdam NL-1081 HV, The Netherlands
3. Department of Biochemistry, University of Stellenbosch, Matieland 7602, South Africa