1. Yunnan Key Laboratory of Plant Reproductive Adaptation and Evolutionary Ecology, and State Key Laboratory for Conservation and Utilization of Bio‐Resources in Yunnan, Institute of Biodiversity, School of Ecology and Environmental Science Yunnan University Kunming 650504 Yunnan China
2. Key Laboratory of Grassland Ecosystem of Ministry of Education, and Sino‐U.S. Centers for Grazingland Ecosystem Sustainability, College of Grassland Science, Gansu Agricultural University Lanzhou 730070 China
3. Kunming Institute of Botany Chinese Academy of Sciences Kunming 650201 China
4. Shangri‐La Alpine Botanical Garden Diqing 674400 China