1. State Key Laboratory of Biocontrol, Institute of Aquatic Economic Animals and the Guangdong Province Key Laboratory for Aquatic Economic Animals, School of Life Sciences; Sun Yat-Sen University; Guangzhou China
2. Marine Fisheries Development Center of Guangdong Province; Huizhou China
3. Key Laboratory of Aquatic Animal Breeding and Culturing for South China Sea; Ministry of Agriculture; Guangzhou China
4. South China Sea Bio-Resource Exploitation and Utilization Collaborative Innovation Center; Guangzhou China