1. Department of Environmental Conservation University of Massachusetts 160 Holdsworth Way Amherst Massachusetts 01003 USA
2. Organismic and Evolutionary Biology Graduate Program University of Massachusetts 204C French Hall, 230 Stockbridge Road Amherst Massachusetts 01003 USA
3. Greater Atlantic Regional Fisheries Office NOAA National Marine Fisheries Service 55 Great Republic Drive Gloucester Massachusetts 01930 USA
4. Centre for Research into Ecological and Environmental Modelling University of St Andrews Fife KY16 9LZ St. Andrews United Kingdom