1. Department of Plant Biology, Ecology& Evolution Oklahoma State University Stillwater OK 74078 USA
2. Department of Biodiversity Earth & Environmental Sciences & Academy of Natural Sciences of Drexel University Philadelphia PA 19103 USA
3. Department of Biology Hobart and William Smith Colleges Geneva NY 14456 USA
4. Departamento de Biologia Vegetal Instituto de Biologia Universidade Estadual de Campinas ‐ UNICAMP CP. 6109 13083‐970 Campinas São Paulo Brazil
5. Department of Biology Bucknell University Lewisburg PA 17837 USA