1. Inra Functional Ecology and Biogeochemistry of Soils and Agrosystems University of Montpellier–CIRAD–INRA–IRD–SupAgro 2 place Pierre Viala Montpellier 34060 France
2. Cirad Functional Ecology and Biogeochemistry of Soils and Agrosystems University of Montpellier–CIRAD–INRA–IRD–SupAgro 2 place Pierre Viala Montpellier 34060 France
3. Department of Ecology, Evolution and Behavior University of Minnesota Twin Cities Saint Paul Minnesota 55108 USA
4. Centre for Biodiversity Theory and Modelling Theoretical and Experimental Ecology Station CNRS and Paul Sabatier University Moulis 09200 France
5. CEFE University of Montpellier–CNRS–EPHE–IRD–University Paul‐Valéry Montpellier 1919 route de Mende Montpellier 34293 France