1. Queen Mary University of London London UK
2. Smithsonian Tropical Research Institute Panama City Panama
3. Zoological Society of London London UK
4. University of Texas at Arlington Arlington Texas USA
5. Lancaster University Lancaster UK
6. Arizona State University Tempe Arizona USA
7. University of Michigan Ann Arbor Michigan USA
8. University of Nevada Reno Nevada USA
9. Princeton University Princeton New Jersey USA
10. Florida International University Miami South Florida USA