1. Department of Ecology and Evolutionary Biology, Yale University, New Haven, Connecticut, USA;
2. Department of Biology, University of Texas at Arlington, Arlington, Texas, USA
3. Department of Ecology and Evolutionary Biology, University of Colorado, Boulder, Colorado, USA
4. Department of Biology, St. Mary's College of Maryland, St. Mary's City, Maryland, USA
5. Department of Ecology and Evolutionary Biology, University of Toronto, Toronto, Ontario, Canada