1. Department of Innovative Medicine Graduate School of Medicine Chiba University Chiba Japan
2. Division of Comparative Pathology and Medicine Department of Pathology University of California San Diego San Diego CA USA
3. Center for Veterinary Sciences and Comparative Medicine University of California San Diego CA USA
4. Departments of Medicine and Pathology CU‐UCSD Center for Mucosal Immunology Allergy and Vaccines (CU‐UCSD cMAV) University of California San Diego CA USA
5. International Research and Development Center for Mucosal Vaccines The Institute of Medical Science The University of Tokyo Tokyo Japan
6. Department of Mucosal Immunology The University of Tokyo Distinguished Professor Unit The Institute of Medical Science The University of Tokyo Tokyo Japan
7. Department of Human Mucosal Vaccinology Chiba University Hospital Chiba Japan